Haplogroup J-M267

Haplogroup J-M267
Possible time of origin	15,000-34,000 years before present
Possible place of origin	Western Asia
Ancestor	J-P209
Descendants	J-M62, J-M365.1, J-L136, J-Z1828
Defining mutations	M267, L255, L321, L765, L814, L827, L1030

Haplogroup J1 redirects here, this page discusses the Y-chromosomal haplogroup of the same name, for the completely separate and distinct mitochondrial haplogroup also named J1 see Haplogroup J (mtDNA)

In Genetic genealogy and human genetics, Y DNA haplogroup J-M267, also commonly known as Haplogroup J1 is a subclade (branch) of Y-DNA haplogroup J-P209, (commonly known as Haplogroup J) along with its sibling clade Y DNA haplogroup J-M172 (commonly known as Haplogroup J2). (All these haplogroups have had other historical names listed below.)

Men from this lineage share a common paternal ancestor, which is demonstrated and defined by the presence of the SNP mutation referred to as M267, which was announced in (Cinnioğlu 2004). This haplogroup is found today in significant frequencies in many areas in or near the Middle East, and parts of the Caucasus, Sudan and Ethiopia. It is also found in high frequencies in parts of North Africa, Southern Europe, and amongst Jewish groups, especially those with Cohen surnames. It can also be found much less commonly, but still occasionally in significant amounts, throughout Europe and as far east as Central Asia and the Indian Subcontinent.

Origins

Since the discovery of haplogroup J-P209 it has generally been recognized that it shows signs of having originated in or near West Asia. The frequency and diversity of both its major branches, J-M267 and J-M172, in that region makes them candidates as genetic markers of the spread of farming technology during the Neolithic, which is proposed to have had a major impact upon human populations.

J-M267 has several recognized subclades, some of which were recognized before J-M267 itself was recognized, for example J-M62 Y Chromosome Consortium "YCC" 2002. With one notable exception, J-P58, most of these are not common (Tofanelli 2009). Because of the dominance of J-P58 in J-M267 populations in many areas, discussion of J-M267's origins require a discussion of J-P58 at the same time.

Distribution

Africa

North Africa and Horn of Africa

North Africa received Semitic migrations, according to some studies it may have been diffused in recent time by Arabs who, mainly from the 7th century a.d., expanded to northern Africa (Arredi 2004 and Semino 2004). However the Canary islands is not known to have had any Semitic language. There J-M267 is dominated by J-P58, and dispersed in a very uneven manner according to studies so far, often but not always being lower among Berber and/or non-urban populations. In Ethiopia there are signs of older movements of J-M267 into Africa across the Red Sea, not only in the J-P58 form. This also appears to be associated with Semitic languages. According to a study in 2011, in Tunisia, J-M267 is significantly more abundant in the urban (31.3%) than in the rural total population (2.5%). According to the authors, these results could be explained by supposing that Arabization in Tunisia was a military enterprise, therefore, mainly driven by men that displaced native Berbers to geographically marginal areas but that frequently married Berber women (Ennafaa 2011).

Population	Sample size	J*(xJ-M172)	total J-M267	J-M267(xP58)	J-P58	publication	previous research on same samples
Algeria (Arabs from Oran)	102	NA	22.5%	NA	NA	Robino 2007
Algeria	20	NA	35%	NA	NA	Semino 2004
Egypt	147	NA	21.1%	1.4%	19.7%	Chiaroni 2009	Luis 2004
Egypt	124	NA	19.8%	NA	NA	El-Sibai 2009
Egypt (Western Desert)	35	NA	31.4%	NA	NA	Kujanová 2009
Libya (Tuareg)	47	NA	0.0%	NA	NA	Ottoni 2011
Libya (Benghazi)	238	NA	39.5%	NA	NA	Alvarez 2014	Elmrghni 2012
Morocco (Amizmiz Valley)	33	NA	0%	NA	NA	Alvarez 2009
Morocco	51	NA	19.6%	NA	NA	Onofri 2008
Morocco (Arabs)	49	NA	10.2%	NA	NA	Semino 2004
Morocco (Arabs)	44	NA	13.6%	NA	NA	Semino 2004
Morocco (Berbers)	64	NA	6.3%	NA	NA	Semino 2004
Morocco (Berbers)	103	NA	7.8%	NA	NA	Semino 2004
Morocco (Rabat)	267	NA	21.3%	NA	NA	Alvarez 2014	Aboukhalid 2010
Morocco (Casablanca)	166	NA	15.7%	NA	NA	Alvarez 2014	Laouina 2011
Morocco (Figuig Oasis)	96	NA	29.2%	NA	NA	Alvarez 2014	Palet 2010
Morocco (El Jadida)	49	NA	8.2%	NA	NA	Alvarez 2014
Morocco (Fes)	108	NA	16.7%	NA	16.7%	Regueiro 2015
Tunisia	73	NA	30.1%	NA	NA	Semino 2004
Tunisia (Sousse)	220	NA	25.9%	NA	25.9%	Fadhlaoui-Zid 2015
Tunisia (Tunis)	148	NA	32.4%	1.3%	31.1%	Grugni 2012	Arredi 2004
Tunisia	52	NA	34.6%	NA	NA	Onofri 2008
Tunisia (Bou Omrane Berbers)	40	NA	0%	NA	NA	Ennafaa 2011
Tunisia (Bou Saad Berbers)	40	NA	5%	0%	5%	Ennafaa 2011
Tunisia (Jerbian Arabs)	46	NA	8.7%	NA	NA	Ennafaa 2011
Tunisia (Jerbian Berbers)	47	NA	0%	NA	NA	Ennafaa 2011
Tunisia (Sened Berbers)	35	NA	31.4%	0%	31.4%	Fadhlaoui-Zid 2011
Tunisia (Andalusian Zaghouan)	32	NA	43.8%	0%	43.8%	Fadhlaoui-Zid 2011
Tunisia (Cosmopolitan Tunis)	33	NA	24.2	0%	24.2%	Fadhlaoui-Zid 2011
Tunisia (Sejenane)	47	NA	34.0%	NA	NA	Alvarez 2014	Frigi 2011
Tunisia (Sfax)	56	NA	25%	NA	25%	Regueiro 2015
Tunisia (Beja)	72	NA	15.3%	NA	15.3%	Regueiro 2015
Canary Islands (pre-Hispanic)	30	NA	16.7%	NA	NA	Fregel 2009
Canary Islands (17th-18thC)	42	NA	11.9%	NA	NA	Fregel 2009
Canary Islands	652	NA	3.5%	NA	NA	Fregel 2009
Sahrawi	89	NA	20.2%	NA	NA	Fregel 2009	Bosch 2001 and Flores 2001
Sudan (Khartoum)	35	NA	74.3%	0.0%	74.3%	Chiaroni 2009	Tofanelli 2009 and Hassan 2008
Sudan-Arabic	35	NA	17.1%	0.0%	17.1%	Chiaroni 2009	Hassan 2008
Sudan (Nilo-Saharan languages)	61	NA	4.9%	3.3%	1.6%	Chiaroni 2009	Hassan 2008
Ethiopia Oromo	78	NA	2.6%	2.6%	0.0%	Chiaroni 2009	Semino 2004
Ethiopia Amhara	48	NA	29.2%	8.3%	20.8%	Chiaroni 2009	Semino 2004
Ethiopia Arsi	85	22%	NA	NA	NA	Moran 2004
Ethiopia General	95	21%	NA	NA	NA	Moran 2004
Comoros Islands	293	NA	5.0%	NA	NA	Msaidie 2011

Asia

South Asia

J*(xJ-M172) was found in India among Indian Muslims.

Population	Sample size	J*(xJ-M172)	total J-M267	J-M267(xP58)	J-P58	Publication
India (Indian Shia)	161	10.6%	NA	NA	NA	Eaaswarkhanth 2009
India (Indian Sunni)	129	2.3%	NA	NA	NA	Eaaswarkhanth 2009
India (Mappla)	40	10%	NA	NA	NA	Eaaswarkhanth 2009

West Asia

The area including eastern Turkey and the Zagros and Taurus mountains, has been identified as a likely area of ancient J-M267 diversity. Both J-P58 and other types of J-M267 are present, sometimes with similar frequencies.

Population	Sample size	Total J-M267	J-M267(xP58)	J-P58	Publication	Previous research on same samples
Turkey	523	9.0%	3.1%	5.9%	Chiaroni 2009	Cinnioğlu 2004
Iran	150	11.3%	2.7%	8.7%	Chiaroni 2009	Regueiro 2006
Kurds Iraq	93	11.8%	4.3%	7.5%	Chiaroni 2009
Assyrians modern Iraq	28	28.6%	17.9%	10.7%	Chiaroni 2009
Iraq (Nassiriya)	56	26.8%	1.8%	25.0%	Chiaroni 2009	Tofanelli 2009
Assyrians Iran	31	16.1%	9.7%	6.5%	Chiaroni 2009
Iran	92	3.2%	NA	NA	El-Sibai 2009
Assyrians Turkey	25	20.0%	16.0%	4.0%	Chiaroni 2009

Levant and Semitic populations

J-M267 is very common throughout this region, dominated by J-P58, but some specific sub-populations have notably low frequencies.

Population	Sample size	Total J-M267	J-M267(xP58)	J-P58	Publication	Previous research on same samples
Syria	554	33.6%	NA	NA	El-Sibai 2009	Zalloua 2008
Druzes (Djebel Druze)	34	14.7%	2.9%	11.8%	Chiaroni 2009
Syria (Sunni from Hama)	36	47.2%	2.8%	44.4%	Chiaroni 2009
Syria (Ma'loula Aramaean)	44	6.8%	4.5%	2.3%	Chiaroni 2009
Syria (Sednaya Syriac Catholic)	14	14.3%	0.0%	14.3%	Chiaroni 2009
Syrian Catholic Damascus	42	9.5%	0.0%	9.5%	Chiaroni 2009
Alawites Syria	45	26.7%	0.0%	26.7%	Chiaroni 2009
Assyrian NE Syria	30	3.3%	0.0%	3.3%	Chiaroni 2009
Ismaili Damascus	51	58.8%	0.0%	58.8%	Chiaroni 2009
Lebanon	951	18.9%	NA	NA	Zalloua 2008
Galilee Druze	172	13.4%	1.2%	12.2%	Chiaroni 2009	Shlush 2008
Palestinians (Akka (Acre))	101	39.2%	NA	NA	Zalloua 2008
Palestine	49	32.7%	0.0%	32.7%	Chiaroni 2009
Jordan	76	48.7%	0.0%	48.7%	Chiaroni 2009
Jordan	273	35.5%	NA	NA	El-Sibai 2009
Jordan (Amman)	101	40.6%	NA	NA	Flores 2005
Jordan (Dead Sea)	45	8.9%	NA	NA	Flores 2005
Jews (Portugal/Trás-os-Montes)	57	12.3%	NA	NA	Nogueiro 2009
Jews (Cohanim)	215	46.0%	0.0%	46.0%	Hammer & Behar 2009
Jews (non Cohanim)	1,360	14.9%	0.9%	14.0%	Hammer 2009
Bedouin Negev	28	67.9%	3.6%	64.3%	Chiaroni 2009	Cann 2002

Arabian peninsula

J-P58 is the most common Y-Chromosome haplogroup among men from all of this region.

Population	Sample size	Total J-M267	J-M267(xP58)	J-P58	Publication	Previous research on same samples
Saudi Arabia	157	40.1%	NA	NA	Abu-Amero 2009
Qatar	72	58.3%	1.4%	56.9%	Chiaroni 2009	Cadenas 2007
UAE	164	34.8%	0.0%	34.8%	Chiaroni 2009	Cadenas 2007
Yemen	62	72.6%	4.8%	67.7%	Chiaroni 2009	Cadenas 2007
Kuwait	42	33.3%	NA	NA	El-Sibai 2009
Oman	121	38.0%	0.8%	37.2%	Chiaroni 2009	Luis 2004

Europe

J-M267 is uncommon in Europe. It is, however, found at levels of 5–10% among many populations in southern Europe.

Population	Sample size	Total J-M267	J-M267(xP58)	J-P58	publication
Malta	90	7.8%	NA	NA	El-Sibai 2009
Crete	193	8.3%	NA	NA	King 2008
Greece (mainland)	171	4.7%	NA	NA	King 2008
Macedonia (Greece)	56	1.8%	NA	NA	Semino 2004
Greece	249	1.6%	NA	NA	Di Giacomo 2004
Bulgaria	808	3.4%	NA	NA	Karachanak 2013
Romania	130	1.5%	NA	NA	Di Giacomo 2004
Russia	223	0.4%	NA	NA	Di Giacomo 2004
Republic of Macedonia Albanian speakers	64	6.3%	NA	NA	Battaglia 2008
Albania	56	3.6%	NA	NA	Semino 2004
Croats (Osijek)	29	0.0%	NA	NA	Battaglia 2008
Slovenia	75	1.3%	NA	NA	Battaglia 2008
Italians (northeast)	67	0.0%	NA	NA	Battaglia 2008
Italians	915	0.7%	NA	NA	Capelli 2009
Sicily	236	3.8%	NA	NA	Di Gaetano 2008
Provence	51	2%	NA	NA	King 2011
Portugal (North)	101	1.0%	NA	NA	Gonçalves 2005
Portugal (Centre)	102	4.9%	NA	NA	Gonçalves 2005
Portugal (South)	100	7.0%	NA	NA	Gonçalves 2005
Açores	121	2.5%	NA	NA	Gonçalves 2005
Madeira	129	0.0%	NA	NA	Gonçalves 2005

Caucasus

The Caucasus has areas of both high and low J-M267 frequency. The J-M267 in the Caucasus is also notable because most of it is not within the J-P58 subclade.

Population	Sample size	Total J-M267	J-M267(xP58)	J-P58	Publication
Avars	115	59.0%	58.0%	1.0%	Balanovsky 2011
Dargins	101	70.0%	69.0%	1.0%	Balanovsky 2011
Kubachi	65	99.0%	99.0%	0.0%	Balanovsky 2011
Kaitak	33	85.0%	85.0%	0.0%	Balanovsky 2011
Lezghins	81	44.4%	44.4%	0.0%	Balanovsky 2011
Shapsug	100	0.0%	0.0%	0.0%	Balanovsky 2011
Abkhaz	58	0.0%	0.0%	0.0%	Balanovsky 2011
Circassians	142	11.9%	4.9%	7.0%	Balanovsky 2011
Ingush	143	2.8%	2.8%	0.0%	Balanovsky 2011
Ossets	357	1.3%	1.3%	0.0%	Balanovsky 2011
Chechens (Ingushetia)	112	21.0%	21.0%	0.0%	Balanovsky 2011
Chechens (Chechnya)	118	25.0%	25.0%	0.0%	Balanovsky 2011
Chechens (Dagestan)	100	16.0%	16.0%	0.0%	Balanovsky 2011
Azerbaijan	46	15.2%	NA	NA	Di Giacomo 2004

Subclade Distribution

J-P58

The P58 marker which defines subgroup J1c3 was announced in (Karafet 2008), but had been announced earlier under the name Page08 in (Repping 2006 and called that again in Chiaroni 2011). It is very prevalent in many areas where J-M267 is common, especially in parts of North Africa and throughout the Arabian peninsula. It also makes up approximately 70% of the J-M267 among the Amhara of Ethiopia. Notably, it is not common among the J-M267.

Chiaroni 2009 proposed that J-P58 (that they refer to as J1e) might have first dispersed during the Pre-Pottery Neolithic B period, "from a geographical zone, including northeast Syria, northern Iraq and eastern Turkey toward Mediterranean Anatolia, Ismaili from southern Syria, Jordan, Palestine and northern Egypt." They further propose that the Zarzian material culture may be ancestral. They also propose that this movement of people may also be linked to the dispersal of Semitic languages by hunter-herders, who moved into arid areas during periods known to have had low rainfall. Thus, while other haplogroups including J-M267 moved out of the area with agriculturalists who followed the rainfall, populations carrying J-M267 remained with their flocks (King 2002 and Chiaroni 2008).

According to this scenario, after the initial neolithic expansion involving Semitic languages, which possibly reached as far as Yemen, a more recent dispersal occurred during the Chalcolithic or Early Bronze Age (approximately 3000–5000 BCE), and this involved the branch of Semitic which leads to the Arabic language. The authors propose that this involved a spread of some J-P58 from the direction of Syria towards Arab populations of the Arabian Peninsula and Negev.

On the other hand, the authors agree that later waves of dispersion in and around this area have also had complex effects upon the distributions of some types of J-P58 in some regions. They list three regions which are particularly important to their proposal:

1. The Levant (Syria, Jordan, Israel and Palestine). In this area, Chiaroni 2009 note a "patchy distribution of J1c3 or J-P58 frequency" which is difficult to interpret, and which "may reflect the complex demographic dynamics of religion and ethnicity in the region".
2. The northern area of eastern Anatolia, northern Iraq and northwest Iran. In this area, Chiaroni 2009 recognize signs that J-M267 might have an older presence, and on balance they accept the evidence but note that it could be in error.
3. The southern area of Oman, Yemen and Ethiopia. In this area, Chiaroni 2009 recognize similar signs, but reject it as possible a result of "either sampling variability and/or demographic complexity associated with multiple founders and multiple migrations."

The "YCAII=22-22 and DYS388≥15" cluster

Not only is the J-P58 group itself very dominant in many areas where J-M267 or J1 is common, but J-P58 in turn contains a large cluster which had been recognized before the discovery of P58, and is still a subject of research. This relatively young cluster, compared to J-M267 overall, was identified by STR markers haplotypes - specifically YCAII as 22-22, and DYS388 having unusual repeat values of 15 or higher, instead of more typical 13 (Chiaroni 2011) This cluster was found to be relevant in some well-publicized studies of Jewish and Palestinian populations (Nebel 2000 and Hammer 2009). More generally, since then this cluster has been found to be frequent among men in the Middle East and North Africa, but less frequent in areas of Ethiopia and Europe where J-M267 is nevertheless common. The pattern is therefore similar to the pattern of J-P58 generally, described above, and may be caused by the same movements of people (Chiaroni 2009).

Tofanelli 2009 refers to this overall cluster with YCAII=22-22 and high DYS388 values as an "Arabic" as opposed to a "Eurasian" type of J-M267. This Arabic type includes Arabic speakers from Maghreb, Sudan, Iraq and Qatar, and it is a relatively homogeneous group, implying that it might have dispersed relatively recently compared to J-M267 generally. The more diverse "Eurasian" group includes Europeans, Kurds, Iranians and Ethiopians (despite Ethiopia being outside of Eurasia), and is much more diverse. The authors also say that "Omanis show a mix of Eurasian pool-like and typical Arabic haplotypes as expected, considering the role of corridor played at different times by the Gulf of Oman in the dispersal of Asian and East African genes." Chiaroni 2009 also noted the anomalously high apparent age of Omani J-M267 when looking more generally at J-P58 and J-M267 more generally.

This cluster in turn contains three well-known related sub-clusters. First, it contains the majority of the Jewish "Cohen modal haplotype", found among Jewish populations, but especially in men with surnames related to Cohen. It also contains both the Galilee modal haplotype and Palestinian & Israeli Arab modal haplotype associated with Palestinians and Israeli Arabs by Nebel 2000 and Hammer 2009. Nebel 2002 then pointed out that the Galilee modal is also the most frequent type of J-P209 haplotype found in northwest Africans, and in Yemen, so it is not isolated to the area of Israel and the Palestine. But notably, this particular variant "is absent from two distinct non-Arab Middle Eastern populations, Jews and Muslim Kurds", even though both these populations do have high levels of J-P209 haplotypes.

Nebel 2002 noted not only the presence of the Galilee modal of J-M267 in the Maghreb but also that J-M267 in this region had very little diversity generally. They concluded that J-M267 in this region "is derived not only from the early Neolithic dispersion but also from recent expansions from the Arabian peninsula" proposing that they might have been carried from the Middle East with the Arab expansion in the seventh century AD. Semino 2004 later agreed that this seemed consistent with the evidence and generalized from this that distribution of the entire YCAII=22-22 cluster of J-M267 in the Arabic speaking areas of the Middle East and North Africa might in fact mainly have an origin in historical times.

More recent studies have emphasized doubt that the Islamic expansions are old enough to completely explain the major patterns of J-M267 frequencies. Chiaroni 2009 rejected this for J-P58 as a whole, but accepted that "some of the populations with low diversity, such as Bedouins from Israel, Qatar, Sudan and UAE, are tightly clustered near high-frequency haplotypes suggesting founder effects with star burst expansion in the Arabian Desert". They did not comment on the Maghreb.

Tofanelli 2009 take a stronger position of rejecting any strong correlation between the Arab expansion and either the YCAII=22-22 STR-defined sub-cluster as discussed by Semino 2004 or the smaller "Galilee modal" as discussed by (Nebel 2002). They also estimate that the Cohen modal haplotype must be older than 4500 years old, and maybe as much as 8600 years old - well before the supposed origin of the Cohanim. Only the so-called Palestinian & Israeli Arab modal had a strong correlation to an ethnic group, but it was also rare. In conclusion, the authors were negative about the usefulness of STR defined modals for any "forensic or genealogical purposes" because "they were found across ethnic groups with different cultural or geographic affiliation".

Hammer 2009 disagreed, at least concerning the Cohen modal haplotype. They said that it was necessary to look at a more detailed STR haplotype in order to define a new "Extended Cohen Modal Haplotype" which is extremely rare outside Jewish populations, and even within Jewish populations is mainly only found in Cohanim. They also said that by using more markers and a more restrictive definition, the estimated age of the Cohanim lineage is lower than the estimates of Tofanelli 2009, and it is consistent with a common ancestor at the approximate time of founding of the priesthood which is the source of Cohen surnames.

J-M368

The correspondence between P58 and high DYS388 values, and YCAII=22-22 is not perfect. For example the J-M267 subclade of J-P58 defined by SNP M368 has DYS388=13 and YCAII=19-22, like other types of J-M267 outside the "Arabic" type of J-M267, and it is therefore believed to be a relatively old offshoot of J-P58, that did not take part in the most recent waves of J-M267 expansion in the Middle East (Chiaroni 2009). These DYS388=13 haplotypes are most common in the Caucasus and Anatolia, but also found in Ethiopia (Tofanelli 2009).

Phylogenetics and Distribution

There are several confirmed and proposed phylogenetic trees available for haplogroup J-M267. The following phylogeny or family tree of J-M267 haplogroup subclades is based on the ISOGG (2012) tree, which is in turn based upon the YCC 2008 tree and subsequent published research.

J1 (L255, L321, M267)

• J1* J1* clusters are found in Eastern Anatolia & parts of the Caucasus.
• J1a (M62) found in a very small frequency in Britain.
• J1b (M365.1) found in a small frequency in Eastern Anatolia, Iran & parts of Europe.
• J1c (L136)

• J1c* Found in a very small frequency in Europe.

• J1c1 (M390)

• J1c2 (P56) found sporadically in Anatolia, East Africa, the Arabian Peninsula & Europe.

• J1c3

• J1c3* found in a low frequency in the Levant & the Arabian Peninsula.

• J1c3a (M367.1, M368.1) - formerly J1e1.

• J1c3b (M369) - formerly J1e2.

• J1c3c (L92, L93) found in a small frequency in South Arabia.

• J1c3d (L147.1) accounts for the majority J1, the predominant haplogroup in Yemen.

• J1c3d* accounts for the majority of J1 in Yemen, Cohen Jews and Ethiopia. as well as Quraysh including Seyyed.

• J1c3d1 (L174.1)

• J1c3d2 (L222.2) Found in the majority of J1c3d in Saudi Arabia & Sudan. An important element of J1c3d in North Africa.

• J1c3d2*

• J1c3d2a (L65.2/S159.2)
• Jl829 found in a part of Idrisid family

See also

Genetics

3

Y-DNA J Subclades

3

Y-DNA Backbone Tree

Evolutionary tree of human Y-chromosome DNA haplogroups
	"Y-chromosomal Adam"
		A00	A0-T
			A0	A1
				A1a	A1b
					A1b1		BT
						B	CT
							DE						CF
						D	E		C		F
															F1	F2		F3	GHIJK
																				G	HIJK
																						H	IJK
																							IJ	K
																						I	J		LT		K2
																										L	T		NO	K2b	K2c	K2d		K2e
																													N	O		K2b1	P
																																	M	S		Q	R
Y-DNA by populations Famous Y-DNA haplotypes

References

Works Cited

Journals

• See also Supplementary Material.

• . (Also see Errata)

Websites

Haplogroups/Phylogeny

• Ongoing Corrections/Additions by citizen scientists.

Haplotype/SNP research Projects. See also Y-DNA haplogroup projects (ISOGG Wiki)

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Haplogroup-Specific Ethnic/Geographical Group Projects

• This is an ongoing research project by citizen scientists. Over 950 J members.
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External links

• J project and J1-M267 project
• J1b Project
• Y-DNA Haplogroup J and its Subclades - 2011